#PING. ALGIDA NAME: [Pinguicula algida] IMAGE: sj70.188.jpg DESC: sj70.188: Winter buds CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: August 29, 1970 TEXT: { P. algida was described as a distinct species by L. Malyshev in 1966. The depicted specimens were collected by himself in the Putorana Mountains (Russia, Siberia, 300 km E of Norilsk, lower Jenissej region, 300-800m) in August 1970. Unfortunately the hibernacula perished as I didn't keep them cold enough for a long time. They need (as also P. variegata) 10-15 centigrades below the freezing point during 9-10 months. } #PING. ALPINA NAME: [Pinguicula alpina] IMAGE: sj72.06.jpg DESC: sj72.06: Three views of flower (natural site specimens) May 22, 1972 IMAGE: sj72.07.jpg DESC: sj72.07: Six flowers (natural site specimens) May 22, 1972 IMAGE: sj78.04.jpg DESC: sj78.04: Habitus (natural site specimens) May 14, 1978 IMAGE: sj78.03.jpg DESC: sj78.03: Habitus (natural site specimens) May 14, 1978 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: TEXT: { Origin of depicted specimens: Switzerland, Canton Bern (border to canton Fribourg), Schwarzwasserbrücke, 600m, calcareous sandstone. Relictic lowland site of the last ice-age, collected by myself. See also pictures with P. leptoceras sj70.187 and P. vulgaris sj72.08. Note the different size of the yellow spot(s) on the central lobe of the lower lip in sj72.07. } NAME: [Pinguicula alpina] IMAGE: sj69.27.jpg DESC: sj69.27: Two flowers (natural site specimens) June 21, 1969 IMAGE: sj69.175.jpg DESC: sj69.175: Winter buds and roots (natural site specimens) October 10, 1969 IMAGE: sj70.148.jpg DESC: sj70.148: Two aberrant flowers (natural site specimens) July 26, 1970 IMAGE: sj70.149.jpg DESC: sj70.149: Two aberrant flowers (natural site specimens) July 26, 1970 IMAGE: sj77.07.jpg DESC: sj77.07: Habitus (natural site specimen) June 17, 1977 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: TEXT: { Origin of depicted specimens: Switzerland, Canton Bern, Prealps, Trogenmoos/Grünenbergpass (N of Interlaken), 1500m, slightly acid soil.
The specimens shown in sj70.148 and sj70.149 have only 4 corolla lobes and the calyx is in axial position at the very end of the tubular corolla throat, 'behind' the rudimentary spur, while in normal flowers it is 'in front' the spur. I guess in this growth disorder the flower development was stopped in an early, 'atavistic' stage, showing how Pinguicula flowers might have looked in an earlier phase of the evolution of this genus.
Pinguicula alpina is the only winter-bud-forming species with perennial roots. For comparison see winter buds of P. algida or P. grandiflora. } NAME: [Pinguicula alpina and other species] IMAGE: sj70.84.jpg DESC: sj70.84: Three flowers (cultivation specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: June 10, 1970 TEXT: { Top: P. alpina, origin see sj.69.27; middle: P. vallisneriifolia, origin see sj70.88; bottom: P. longifolia subsp. reichenbachiana, origin see sj70.70 } NAME: [Pinguicula alpina] [Pinguicula vulgaris] IMAGE: sj77.27.jpg DESC: sj77.27: Leaves (natural site specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 22, 1977 TEXT: { Common origin of depicted specimens: see P. alpina sj69.27. Left: P. vulgaris, right: P. alpina. At common sites the two species can be distinguished in the non-flowering stage as follows: 1) P. alpina has strong, yellowish, perennial, roots, while the softer, whitish roots of P. vulgaris decay in winter. 2) The leaves of P. alpina are narrower and lack digestive glands over the mid-rib, while the larger leaves of P. vulgaris have a regular glandular structure all over their surface. P. alpinaflowers 2-4 weeks earlier than P. vulgaris. } #PING. BALCANICA NAME: [Pinguicula balcanica] IMAGE: sj77.13.jpg DESC: sj77.13: Three views of flower (cultivation specimens) July 16, 1977 IMAGE: sj78.35.jpg DESC: sj78.35: Habitus (cultivation specimens) July 23, 1978 IMAGE: sj81.25.jpg DESC: sj81.25: Leaves (cultivation specimens) July 23, 1981 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} TEXT: { Origin of depicted specimens: Bulgaria, Vitosha Mt. (south of Sofia) between Stastliveca Hotel and the summit of Cerni Vrach, 2200m, granite, collected by myself September 17, 1976. See also P. vulgaris)
Note the numerous little flies sitting on the flowers of these cultivation specimens in photo sj78.35.
Even in sites with full sun, the specimens of the Vitosha site display a conspicuous difference of pigmentation of the leaves. Some rosettes remain absolutely green while others develop brownish-reddish pigmentation. This difference also remains in cultivation as shown in picture sj81.25. The flowers of both leaf types do not differ in color. } NAME: [Pinguicula balcanica] IMAGE: sj95.9.17.jpg DESC: sj95.9.17: Habitat (natural site) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 26, 1995 TEXT: { Origin of depicted specimens: Central Greece, Vardoussia Mt., Pittimaliko above Anastasios Diakos, between Stavros and upper lodge, 1800-1900m, limestone, site visited by Jan Schlauer and myself July 26, 1995. This is up to now the southernmost known site of P. balcanica. In difference of part of the plants of the Vitosha Mt. (Bulgaria) the leaves of the Vardoussia populations have no reddish pigmentation at all. } NAME: [Pinguicula balcanica] IMAGE: sj95.10.10.jpg DESC: sj95.10.10: Habitat CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 27,1995 TEXT: { Origin of depicted specimens: Greece, Epirus, Smolikas Mt., above Agia Paraskevi, 1700-1850m, serpentine. As the plants from the Vardoussia Mt. and in difference to the bulgarian Vitosha Mt. plants the leaves of this site lack any reddish pigmentation even in full sun. } #PING. CORSICA NAME: [Pinguicula corsica] IMAGE: sj70.153.jpg DESC: sj70.153: Three views of flower (cultivation specimens) July 27, 1970 IMAGE: sj70.111.jpg DESC: sj70.111: Habitus (cultivation specimen) July 5, 1970 IMAGE: sj72.25.jpg DESC: sj72.25: Habitus (cultivation specimen) June 25, 1972 IMAGE: sj79.35.jpg DESC: sj79.35: Habitus (cultivation specimen) March 26, 1979 IMAGE: sj80.37.jpg DESC: sj80.37: 2 flowers (cultivation specimen) June 1, 1980 IMAGE: sj89.1.31.jpg DESC: sj89.1.31: Habitus (cultivation specimen) June 26, 1989 IMAGE: sj82.10.jpg DESC: sj82.10: Habitat IMAGE: sj70.210.jpg DESC: sj70.210: Habitat IMAGE: sj82.05.jpg DESC: sj82.05: Two flowers (color variation, cultivation specimens) June 14, 1982 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: TEXT: { Endemic species of the Corsica island. Origin of sj70.111, sj70.153, sj72.25, sj79.35, sj80.37, sj89.1.31: France, central Corsica, Mt. Rotondo, above Lac de Melo, 1700m, granite. Image sj70.210 shows this area. At this site the majority of P. corsicaflowers are almost white. Plants collected by Mrs. M. Conrad on August 24, 1969. (Pictures taken later in cultivation.
The white flower in sj82.05 was also collected from the same location as sj70.153. The lilac flower was collected from France, central Corsica, Mt. Rotondo, Lac d'Oriente, 2060m, granite. Plants collected by Mrs. M. Conrad on Sept. 4, 1979. The flower color of Pinguicula corsica is known to be extremely variable from site to site.
sj82.10 shows a typical habitat in France, central Corsica, Lac de Nino between and below Mt. Tozzo and Mt. Cimatella, 1750 m, granite. At this site most P. corsicaflowers are lilac-pinkish.
The plants I still grow today (1996) are all offsprings of the few specimens received by Mrs. Conrad in 1969 and 1979, respectively.
} #PING. CRYSTALLINA NAME: [Pinguicula crystallina] IMAGE: sj80.13.jpg DESC: sj80.13: Three views of flower (natural site specimens) IMAGE: sj80.84.jpg DESC: sj80.84: 3 flowers (cultivation specimens) August 4, 1980 IMAGE: sj80.83.jpg DESC: sj80.83: Habitus (cultivation specimen) August 4, 1980 IMAGE: sj80.19.jpg DESC: sj80.19: Habitat, May 21, 1980 IMAGE: sj80.25.jpg DESC: sj80.25: Habitat, May 21, 1980 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: May 26, 1980 TEXT: { Origin of depicted specimens: Cyprus, south of Troodos (Mt. Olympus) summit, above Pano Platres, 1360 m, limestone, collected May 21, 1980 by myself. At this very small site the flowers of P. crystallina are white. It is likely that the 'true' P. crystallina is an endemism of Cyprus. The closely related populations found in southwestern Turkey are listed here under P. crystallina subsp. hirtiflora. Morphologically and caryologically the P. crystallina/hirtiflora-group is the most interesting Pinguicula species in Europe.
In order to collect a few P. crystallina it was necessary to construct a rope ladder. } NAME: [Pinguicula crystallina] IMAGE: sj94.5.10.jpg DESC: sj94.5.10: Habitus (cultivation specimens), May 1st, 1994 IMAGE: sj94.6.21.jpg DESC: sj94.6.21: Habitus (cultivation specimens), May 9, 1994 IMAGE: khidalgo93.1.jpg DESC: khidalgo93.1: Habitat, October 27, 1993 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} TEXT: { Origin of depicted specimen: Cyprus, N of Troodos (Mt. Olympus) summit, near Ayios Nikolaos tis Steyis Monastery, above Kakopetria, 800m, limestone, collected October 27, 1993, by Mrs. Kathy Hidalgo (see khidalgo93.1). At this site the flowers, mainly the three lower lobes, have a lilac margin. Note the two comparingly small upper corolla lobes. } #PING. CRYSTALLINA SUBSP. HIRTIFLORA NAME: [Pinguicula crystallina subsp. hirtiflora] IMAGE: sj76.43.jpg DESC: sj76.43: Three views of flower (natural site specimens), July 2, 1976 IMAGE: sj76.40.jpg DESC: sj76.40: Habitus (natural site), June 29, 1976 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 2, 1976 TEXT: { Origin of depicted specimens: Italy, Campania, Valle delle Ferriere above Amalfi, 200m, limestone, collected June 29, 1976, by myself. This subspecies is considered by S.J. Casper and several other authors as a distinct species (P. hirtiflora). As also P. crystallina it varies morphologically from site to site. There is a gradient of morphological and caryological characteristics from Italy through Ex-Yugoslavia, Albania; Greece and Turkey till Cyprus. J. Contandriopoulos identified within this subspecies 4 ploidy groups (2n=16, 24, 32 and 48) } NAME: [Pinguicula crystallina subsp. hirtiflora] IMAGE: sj94.2.33A.jpg DESC: sj94.2.33A: Three views of flower (cultivation specimens), April 4, 1994 IMAGE: sj93.7.3.jpg DESC: sj93.7.3: Habitus (cultivation specimen), November 7, 1993 IMAGE: sj94.5.3.jpg DESC: sj94.5.3: Habitus (cultivation specimens), May 1st, 1994 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} TEXT: { Origin of depicted specimens: Italy, Campania, Vietri sul Mare, at sea level, limestone, collected August 2, 1993, by myself. Note the relatively wide two upper corolla lobes. } NAME: [Pinguicula crystallina subsp. hirtiflora] IMAGE: sj94.3.4A.jpg DESC: sj94.3.4A: Three views of flower (cultivation specimens), April 11, 1994 IMAGE: sj84.14.jpg DESC: sj84.14: Habitus (cultivation specimens), June 3, 1984 IMAGE: sj84.12.jpg DESC: sj84.12: Habitus (cultivation specimens), June 3, 1984 IMAGE: sj84.13.jpg DESC: sj84.13: Habitus (cultivation specimens), June 3, 1984 IMAGE: sj93.7.5.jpg DESC: sj93.7.5: Habitus (cultivation specimen), November 7, 1993 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} TEXT: { Origin of depicted specimens: Italy, Calabria, Celadi Gorge below Rossano, 100 m, reddish limestone, collected July 30, 1993, by myself. Note the relatively upper two corolla lobes.
sj84.12-sj84.14 were collected by Otfried Paulsen in spring 1984. } NAME: [Pinguicula crystallina subsp. hirtiflora] IMAGE: sj94.3.17A.jpg DESC: sj94.3.17A: Three views of flower (cultivation specimens), April 11, 1994 IMAGE: sj93.5.36.jpg DESC: sj93.5.36: Flower color variations at natural site, October 23, 1993 IMAGE: sj93.5.17.jpg DESC: sj93.5.17: Habitat, October 23, 1993 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: April 11, 1994 TEXT: { Origin of depicted specimens: SW Turkey, gorge S of Salda Gölü Lake (between Denizli and Burdur), 1250 m, collected October 23, 1993, by myself. At this site about 20 percent of the flowers are pure white.
P. cryst. subsp. hirtifl. grows on both sides of the brooklet in sj93.5.17 mostly with Parnassia palustris . } NAME: [Pinguicula crystallina subsp. hirtiflora] IMAGE: sj94.3.21A.jpg DESC: sj94.3.21A: Three views of flower (cultivation specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: April 11, 1994 TEXT: { Origin of depicted specimens: SW Turkey, gorge between Mugla and Göktepe, several sites, approx. 1000 m, limestone, collected October 24, 1994, by myself. At this site all corolla lobes are relatively narrow. } NAME: [Pinguicula crystallina subsp. hirtiflora] IMAGE: sj95.14.8A.jpg DESC: sj95.14.8A: Flowers (cultivation specimens), September 27, 1995 IMAGE: sj95.11.3.jpg DESC: sj95.11.3: Habitus (natural site), July 31, 1995 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: September 27, 1995 TEXT: { Origin of depicted specimens: Greece, Thessalia, Mt. Olympus, gorge of Lithochoron, 900-1100 m, limestone, collected by Jan Schlauer and myself July 30, 1995. } NAME: [Pinguicula crystallina subsp. hirtiflora] IMAGE: sj95.14.6A.jpg DESC: sj95.14.6A: Flowers (cultivation specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: September 27, 1995 TEXT: { Origin of depicted specimens: Greece, Epirus, Smolikas Mt., above Agia Paraskevi, 1300 m, limestone, collected by Jan Schlauer and myself July 27, 1995. } NAME: [Pinguicula crystallina subsp. hirtiflora f. pallida] IMAGE: sj95.9.10.jpg DESC: sj95.9.10: Habitus (natural site) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 26, 1995 TEXT: { Origin of depicted specimens: Central Greece, Vardoussia Mt., between Athanassios Diakos and the Pittimaliko lodges, 1700 m, limestone, collected by Jan Schlauer and myself July 26, 1995. About 25 percent of the plants have white flowers (f. pallida). } NAME: [Pinguicula crystallina subsp. hirtiflora var. gionae] IMAGE: sj95.14.18A.jpg DESC: sj95.14.18A: Habitus (natural site) IMAGE: sj95.14.24A.jpg DESC: sj95.14.24A: Habitat, October 1st, 1995 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: October 1st, 1995 TEXT: { Origin of depicted specimens: Central Greece, Ghiona Mt., Rekka-Valley, 1300 m, limestone, collected by Ivo Koudela and myself October 1st, 1995. } NAME: [Pinguicula crystallina and subsp. hirtiflora] IMAGE: sj94.3.34A.jpg DESC: sj94.3.34A: Habitus (cultivation specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: April 11, 1994 TEXT: { Origin of depicted specimens: Left: P. cryst. ssp. hirtiflora from Vietri (origin see sj 94.2.33a). Right top: P. cryst. ssp. hirtiflora from Mugla (origin see sj93.3.21A). Right bottom: P. crystallina from Cyprus (origin sse sj94.5.10). } #PING. FIORII NAME: [Pinguicula fiorii] IMAGE: sj94.9.3.8.jpg DESC: sj94.9.3.8: 2 flowers (cultivation specimens), May 28, 1994 IMAGE: sj94.6.30.jpg DESC: sj94.6.30: Habitus (cultivation specimens), May 9, 1994 IMAGE: sj94.8.10.jpg DESC: sj94.8.10: Habitus (cultivation specimens), May 23, 1994 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} TEXT: { Origin of depicted specimens: Italy, Abruzzi, Maiella Mt., Bocca di Valle W of Guardiagrele, 750 m, limestone, collected by myself July 31, 1993. This is the typus site. The taxonomical classification and geographical distribution of this newly detected species isn't yet definitely cleared. Note the P. vulgaris-like leaves and the large and strong calyx which distinguishes this species from P. longifolia subsp. reichenbachiana erroneously indicated from the Abruzzi region by some authors. } #PING. GRANDIFLORA NAME: [Pinguicula grandiflora] IMAGE: sj72.17.jpg DESC: sj72.17: Three views of flower (cultivation specimens), June 4, 1972 IMAGE: sj70.91.jpg DESC: sj70.91: Two flowers (cultivation specimens), June 10, 1970 IMAGE: sj94.8.32.jpg DESC: sj94.8.32: Flowers (cultivation specimens), May 25, 1994 IMAGE: sj72.10.jpg DESC: sj72.10: Habitus (cultivation specimen), June 3, 1972 IMAGE: sj79.34.jpg DESC: sj79.34: Habitus (cultivation specimen), May 26, 1979 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} TEXT: { Origin of depicted specimens: France, Dépt. Ain, near border to Dépt. Jura. between Col de la Faucille and Mijoux (20 km N of the Swiss city Geneva), 1200 m, loamy limestone soil, collected by myself May 28, 1970. At this site as in some others of this region the dark flowered 'normal' form is associated with the f. pallida. See also sj70.105 with P.grandifl. subsp. rosea.
sj70.91 shows on the left a specimen from sunny place, on the right is a specimen from shady place, or hybrid between P. grandifl. and P. grandifl. f. pallida which are both found at this site. } NAME: [Pinguicula grandiflora] IMAGE: sj56.02.jpg DESC: sj56.02: Flower CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: May 30, 1956 TEXT: { Origin of depicted specimens: Ireland, Clare county, The Burren near Ballyvaughan, 400 m, limestone, collected May 1956 by D.A. Webb. The corolla lobes of the Irish and Pyreneen specimens are lesser overlapping than those from the Jura. See also steiger56.03 P. grandifl. f. chionopetra. } NAME: [Pinguicula grandiflora] IMAGE: sj72.30.jpg DESC: sj72.30: Two flowers (cultivation specimens), June 30, 1972 IMAGE: sj79.36.jpg DESC: sj79.36: Habitus (cultivation specimen), May 26, 1979 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} TEXT: { Left: Specimen from the Jura (see sj72.17). Right: Specimen from France, Dépt. Hautes-Pyrénées, between Gèdre and Gavarnie, 1200 m, granite, collected by myself August 22, 1969. The Pyreneen as well as the Irish (see sj56.02) specimens have lesser overlapping corolla lobes than those from the Jura. } NAME: [Pinguicula grandiflora and other species] IMAGE: sj70.132.jpg DESC: sj70.132: Eight flowers (cultivation specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 11, 1970 TEXT: { From left to right: Top: Pinguicula grandiflora, P. grandiflora f. pallida, P. leptoceras, P. villosa. Bottom: P. vulgaris, P. vulgaris f. bicolor, P. corsica, P. macroceras. } NAME: [Pinguicula grandiflora] IMAGE: sj94.4.15A.jpg DESC: sj94.4.15A: Leaves with prey. CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: April 28, 1994 TEXT: { Specimen with captured crane fly. It is questionable whether the digestive enzymes can penetrate the chitin integument of such large prey. After about a week the (undigested?) prey got infected by fungi. See also sj94.4.13A P. grandifl. f. pallida } NAME: [Pinguicula grandiflora, f. pallida and subsp. rosea] IMAGE: sj70.105.jpg DESC: sj70.105: Three flowers CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: June 24, 1970 TEXT: { From left to right: P. grandiflora, P. grandifl. f. pallida, P. grandifl. subsp. rosea } NAME: [Pinguicula grandiflora and P. grandifl. f. chionopetra] IMAGE: sj56.03.jpg DESC: sj56.03: Habitus CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: May 30, 1956 TEXT: { Origin of depicted specimens: Ireland, Clare county, The Burren near Ballyvaughan, 400 m, limestone, collected May 1956 by D.A. Webb. The corolla lobes of the Irish and Pyreneen specimens are lesser overlapping than those from the Jura. See also sj56.02 P. grandiflora. } #PING. GRANDIFLORA F. PALLIDA NAME: [Pinguicula grandiflora f. pallida] IMAGE: sj70.78.jpg DESC: sj70.78: Three views of flower (cultivation specimens), June 10, 1970 IMAGE: sj94.8.21.jpg DESC: sj94.8.21: Flowers (cultivation specimens), May 23, 1994 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} TEXT: { Origin of depicted specimens: France, Dépt. Ain, near border to Dépt. Jura, between Col de la Faucille and Mijoux (20 km N of the Swiss city Geneva), 1200 m, loamy limestone soil, collected by myself May 28, 1970. At this site as in some others of this region the f. pallida is associated with the dark flowered 'normal' form. There is also at least one site where exclusively the f. pallida is found. See also sj70.144 (with P. longifolia ssp. longifolia) and sj70.132 (with other species). } NAME: [Pinguicula grandiflora f. pallida and subsp. rosea] IMAGE: sj70.107.jpg DESC: sj70.107: Flowers (cultivation specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: June 24, 1970 TEXT: { Left: P. grandifl. f. pallida. Right: P. grandifl. subsp. rosea. Note the somewhat shorter spur of the subsp. rosea. } NAME: [Pinguicula grandiflora f. pallida] IMAGE: sj72.46.jpg DESC: sj72.46: Habitus (cultivation specimens), July 9, 1972 IMAGE: sj72.47.jpg DESC: sj72.47: Habitus (cultivation specimens), July 9, 1972 IMAGE: sj75.32.jpg DESC: sj75.32: Habitus (cultivation specimen), June 29, 1975 IMAGE: sj93.2.16.jpg DESC: sj93.2.16: Habitus (cultivation specimen), June 6, 1993 IMAGE: sj94.8.29.jpg DESC: sj94.8.29: Typical flower and f. pallida (cultivation specimens), May 5, 1994 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 9, 1972 TEXT: { Origin of depicted specimens: France, Dépt. Ain, near border to Dépt. Jura. between Col de la Faucille and Mijoux (20 km N of the Swiss city Geneva), 1200 m, loamy limestone soil, collected by myself May 28, 1970. At this site as in some others of this region the dark flowered 'normal' form is associated with the f. pallida. See also sj70.105 with P.grandifl. subsp. rosea. } NAME: [Pinguicula grandiflora f. pallida] IMAGE: sj94.4.13A.jpg DESC: sj94.4.13A: Leaves with prey (cultivation specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: April 28, 1994 TEXT: { Specimen with captured crane fly. It is questionable whether the digestive enzymes can penetrate the chitin integument of such large prey. After about a week the (undigested?) prey got infected by fungi. See also sj94.4.15A P. grandiflora } NAME: [Pinguicula grandiflora f. pallida and Pinguicula grandiflora] IMAGE: sj75.05.jpg DESC: sj75.05: 2 winter buds, gemmae (cultivation specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: January 2, 1975 TEXT: { Left: P. grandifl. f. pallida. Right: P. grandiflora with gemmae which were loosened while preparing the bud for photography. Large hibernacula may produce up to 30 gemmae. } NAME: [Pinguicula grandiflora f. pallida and subsp. rosea] IMAGE: sj95.16.11A.jpg DESC: sj95.16.11A: 2 winter buds, gemmae (cultivation specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: October 22, 1995 TEXT: { Left: P. grandifl. f. pallida. Right P. grandifl. subsp. rosea. } #PING. GRANDIFLORA SUBSP. ROSEA NAME: [Pinguicula grandiflora subsp. rosea] IMAGE: sj70.77.jpg DESC: sj70.77: Three views of flower (cultivation specimens), June 10, 1970 IMAGE: sj94.8.23.jpg DESC: sj94.8.23: Flowers (cultivation specimens), May 23, 1994 IMAGE: sj93.2.2.jpg DESC: sj93.2.2: Habitus (natural site), May 19, 1993 IMAGE: sj72.12.jpg DESC: sj72.12: Habitus (cultivation specimen), June 4, 1972 IMAGE: sj72.37.jpg DESC: sj72.37: Habitus (cultivation specimen), July 7, 1972 IMAGE: sj94.7.10.jpg DESC: sj94.7.10: Habitus (cultivation specimen), May 21, 1994 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} TEXT: { Origin of depicted specimens: France, Dépt. Isère, between Goncelin et Sollières, NE of Grenoble, 700 m, loamy limestone soil, collected by myself May 28, 1970. At the sites of P. grandifl. subsp. rosea the 'normal' P. grandiflora is not found. The spur of P. grandifl. subsp. rosea is 1-3 mm shorter than the spur of P. grandiflora and f. pallida. } #PING. GYPSICOLA NAME: [Pinguicula gypsicola] IMAGE: sj70.194.jpg DESC: sj70.194: (cultivation specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: TEXT: { Origin of depicted specimens: University of Bern Botanical Garden. True origin unknown. } #PING. LEPTOCERAS NAME: [Pinguicula leptoceras] IMAGE: sj70.127.jpg DESC: sj70.127: Flowers (cultivation specimens from typus site), July 7, 1970 IMAGE: sj94.6.36.jpg DESC: sj94.6.36: Flowers (cultivation specimens), May 9, 1994 IMAGE: sj81.29.jpg DESC: sj81.29: Habitus (natural site, typus site), August 12, 1981 IMAGE: sj81.16.jpg DESC: sj81.16: Habitus (cultivation specimens), May 17, 1981 IMAGE: sj95.01.jpg DESC: sj95.01: Habitat (typus site) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 7, 1970 TEXT: { Origin of depicted specimens: Switzerland, canton Bern, Grimsel-Pass, 1800 m, granite (site of the holotypus), collected by myself August 16, 1968. sj95.01 shows the region of the typus site in summer. At left Grimsel-Pass (highest point at the little lake, 2165 m). P. leptoceras grows on both sides below the pass. On the Pinguicula-sites the snows lasts for 8-9 months each year.
The latin name leptoceras is misleading: Usually the spur isn't slender at all. See also pictures sj70.132 (with P. grandiflora and other sp.) and sj70.146 (with P. longifolia and other sp.). } NAME: [Pinguicula leptoceras] IMAGE: sj70.175.jpg DESC: sj70.175: Three views of flower (dark form) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: August 7, 1970 TEXT: { Origin of depicted specimens: Switzerland, canton Graubünden, Engadin, Bernina-Pass, 2300 m, collected by myself August 4, 1970. In the Engadin region specimens which lack any white spots in the flower are not rare. } NAME: [Pinguicula leptoceras] IMAGE: sj70.126.jpg DESC: sj70.126: Three views of flower (whitish form) IMAGE: sj94.6.15.jpg DESC: sj94.6.15: Habitus (cultivation specimens), May 9, 1994 IMAGE: sj93.2.23.jpg DESC: sj93.2.23: Habitus (cultivation specimens), June 6, 1993 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 11, 1970 TEXT: { Origin of depicted specimens: Italy, Piedmont, Maritime Alps, between Colle di Tenda and Cime de Salante (at Italian-French border), 1980 m, limestone, collected by myself September 8, 1968. At this site the corolla lobes are almost white. See also sj70.154.
At the right margin of sj94.6.15, a flower bud of P. grandiflora subsp. rosea can be seen. } NAME: [Pinguicula leptoceras] IMAGE: sj70.154.jpg DESC: sj70.154: Eight flowers of two sites (cultivation specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 27, 1970 TEXT: { Top: Specimens from Switzerland, Grimsel-Pass (see sj70.127). Bottom: Specimens from the Maritime Alps (see sj70.126) } NAME: [Pinguicula leptoceras and other species] IMAGE: sj70.184.jpg DESC: sj70.184: Four flowers (cultivation specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 31, 1970 TEXT: { From left to right: P. leptoceras, P. nevadensis, P. vulgaris, P. alpina } #PING. LONGIFOLIA SUBSP. CAUSSENSIS NAME: [Pinguicula longifolia subsp. caussensis] IMAGE: sj70.73.jpg DESC: sj70.73: Three views of flower (cultivation specimen) IMAGE: sj72.03.jpg DESC: sj72.03: Habitus (cultivation specimens), May 22, 1972 IMAGE: sj72.13.jpg DESC: sj72.13: Habitus (cultivation specimens) June 4, 1972 IMAGE: sj78.15.jpg DESC: sj78.15: Habitat CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: June 8, 1970 TEXT: { Origin of depicted specimens: France, Dépt. Lozère, Gorges du Tarn, 400-520m, limestone (see sj78.15), collected by myself September 4, 1968.
The petrophilous P. longif. subsp. caussensis grows on vertical or overhanging limestone rock walls at places which are not exposed to direct rainfall.
The adjective caussensis is derived from the local mountain range named 'Les Causses'. Flowers see also sj70.86 (with P. longif. subsp. reichenbachiana). } #PING. LONGIFOLIA SUBSP. DERTOSENSIS NAME: [Pinguicula longifolia subsp. dertosensis] IMAGE: sj95.6.9A.jpg DESC: sj95.6.9A: Habitus (natural site), April 18, 1995 IMAGE: sj95.6.18A.jpg DESC: sj95.6.18A: Habitus (natural site), April 18, 1995 IMAGE: sj95.6.3A.jpg DESC: sj95.6.3A: Habitat, April 18, 1995 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} TEXT: { Origin of depicted specimens: Spain, Prov. de Tarragona, Ports de Beseit W of Tortosa, Mt. Caro/Sierra de la Fortalesa, 500-600 m, limestone (see sj95.6.3A), collected by myself April 18, 1995. Up to now the warmest known site for P. longifolia. } NAME: [Pinguicula longifolia subsp. dertosensis] IMAGE: sj95.2.14.jpg DESC: sj95.2.14: Habitus (natural site), April 15, 1995 IMAGE: sj95.2.32.jpg DESC: sj95.2.32: Habitus (natural site), April 15, 1995 IMAGE: sj95.2.37.jpg DESC: sj95.2.37: Habitat, April 15, 1995 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: April 15, 1995 TEXT: { Origin of depicted specimens: Spain, Prov. de Cuenca, northernmost part of the Sierra de Cuenca, Rio Guadiela, Hoz de Beteta, 1200 m, limestone.
It is astonishing how this species could survive in these relatively dry rocks for thousands of years.
I think the P. longifolia-group of this site is more or less identical with the subspecies from the Tortosa region and therefore I list it among P. longif. subsp. dertosensis. } NAME: [Pinguicula longifolia subsp. dertosensis] IMAGE: sj95.3.2.jpg DESC: sj95.3.2: Habitat CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: April 15, 1995 TEXT: { Another habitat in that region: Spain, Prov. de Cuenca, northernmost part of the Sierra de Cuenca, Rio Escabas, bridge between Priego and Cañamares, 870 m, limestone, visited April 15, 1995. The specimens of this site are morphologically identical to those of Hoz de Beteta, but with less color variation and generally darker. } #PING. LONGIFOLIA SUBSP. LONGIFOLIA NAME: [Pinguicula longifolia subsp. longifolia] IMAGE: sj70.144.jpg DESC: sj70.144: Three views of flower (cultivation specimen), July 28, 1970 IMAGE: sj69.143.jpg DESC: sj69.143: Habitat IMAGE: sj72.42.jpg DESC: sj72.42: Habitus (cultivation specimens), July 9, 1972 IMAGE: sj79.29.jpg DESC: sj79.29: Habitus (cultivation specimens), May 26, 1979 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 28,1970 TEXT: { Origin of depicted specimens: France, Dépt. Hautes-Pyrénées, below Cirque du Gavarnie, 1600 m, limestone, in association with Pinguicula alpina and at some places with Pinguicula grandiflora (with which it hybridizes).
This region is part of a national park where collecting flowers without permission is forbidden. } NAME: [Pinguicula longifolia subsp. longifolia and other sp.] IMAGE: sj70.155.jpg DESC: sj70.155: Three flowers (cultivation specimens) July 27, 1970 IMAGE: sj70.146.jpg DESC: sj70.146: Four flowers (cultivation specimens) July 28, 1970 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 27,1970 TEXT: { From left to right in sj70.155: P. longif subsp. longifolia, P. corsica, P. grandifl. f. pallida.
From left to right in sj70.146: P. longif.subsp. longifolia, P.leptoceras (Maritime Alps type), P. vulgaris f. bicolor, P. vulgaris. } #PING. LONGIFOLIA SUBSP. REICHENBACHIANA NAME: [Pinguicula longifolia subsp. reichenbachiana] IMAGE: sj70.70.jpg DESC: sj70.70: Three views of flower (cultivation specimens) IMAGE: sj76.15.jpg DESC: sj76.15: Color variation of cultivated flowers, April 19, 1976 IMAGE: sj94.5.13.jpg DESC: sj94.5.13: Habitus (cultivation specimens), May 1st, 1994 IMAGE: sj94.5.14.jpg DESC: sj94.5.14: Habitus (cultivation specimens), May 1st, 1994 IMAGE: sj69.14.jpg DESC: sj69.14: Habitus (cultivation specimens), May 11, 1969 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: June 8, 1970 TEXT: { Origin of depicted specimens: France, Dépt. Alpes-Maritimes, Maritime Alps, Roya Valley, Gorges de Bergue, 450-600 m, limestone, collected by myself September 8, 1968. See also sj70.84 (with P. alpina and other species).
The flower colors vary remarkably from violet/white to almost pure white. } NAME: [Pinguicula longifolia subsp. reichenbachiana and caussensis] IMAGE: sj70.86.jpg DESC: sj70.86: four flowers (cultivation specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: June 10,1970 TEXT: { 2 larger flowers: P. longif. subsp. caussensisr , 2 smaller flowers: P. longif. subsp. reichenbachiana } #PING. LONGIFOLIA 'MUNDO-TYPE' NAME: [Pinguicula longifolia 'Mundo'-type] IMAGE: sj95.7.30.jpg DESC: sj95.7.30: Flowers (cultivation specimens) IMAGE: sj95.7.31.jpg DESC: sj95.7.31: Flowers (cultivation specimens) IMAGE: sj95.3.13.jpg DESC: sj95.3.13: Habitat, April 15, 1995 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: June 4, 1995 TEXT: { Origin of depicted specimens: Spain, Prov. de Albacete, between Alcaraz and Orcera, Nascimiente del Rio Mundo/Cueva de los Chorros, 950-1100 m, limestone, collected by myself April 15, 1995. See sj95.3.13. This 'species' is something in between P. longifolia and P. vallisneriifolia. Geographically and climatically the evolution of sort of a transition stage between both species could be plausible. The leaves are more longifolia-like whereas the corolla, particularly the calyx and spur, tend more towards P. vallisneriifolia. } #PING. LUSITANICA NAME: [Pinguicula lusitanica] IMAGE: sj75.40.jpg DESC: sj75.40: Three views of flower (cultivation specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: August 11, 1975 TEXT: { Origin of depicted specimens: Ireland, Clare county, Lake Goller between Ennistimon and Lisdoonvarna, 320 m, peat-bog, collected by myself May 31, 1975. } #PING. MACROCERAS NAME: [Pinguicula macroceras] IMAGE: sj70.135.jpg DESC: sj70.135: Three views of flower (cultivation specimens), July 12, 1970 IMAGE: sj70.118.jpg DESC: sj70.118: Habitus (cultivation specimens), July 11, 1970 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 12, 1970 TEXT: { Origin of depicted specimens: Japan, Honshu, Tochigi Pref., below Mt. Nantai-san, site altitude not recorded, volcanic substrate, collected by Masahiro Kondo (late father of Katsuhiko Kondo) and sent to me October 11, 1968. Habitat not identical with the Mt. Nanai-san habitat of P. ramosa. Note the unique, speckled flower. All specimens from that site were speckled. See also sj70.135. } NAME: [Pinguicula macroceras] IMAGE: sj71.26.jpg DESC: sj71.26: Three views of flower (natural site specimens), August 30, 1971 IMAGE: sj71.27.jpg DESC: sj71.27: Three views of flower (natural site specimens), August 30, 1971 IMAGE: sj71.24.jpg DESC: sj71.24: Habitus (natural site), August 29, 1971 IMAGE: sj71.31.jpg DESC: sj71.31: 15 calyces (natural site specimens), August 29, 1971 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: August 30, 1971 TEXT: { Origin of depicted specimens: USA, Washington, Whatcom county, Mt. Baker Lodge, upper Bagley Lake, 1600 m, collected by myself August 29, 1971.
Compare sj71.31 with sj71.18 (P. macroc. subsp. nortensis). Note the narrower, pointed calyx tips. } NAME: [Pinguicula macroceras] IMAGE: sj74.41.jpg DESC: sj74.41: Three flowers (cultivation specimens), June 7, 1974 IMAGE: sj74.57.jpg DESC: sj74.57: Habitus (cultivation specimen), August 10, 1974 IMAGE: sj74.53.jpg DESC: sj74.53: Habitus (cultivation specimens), July 11, 1970 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} TEXT: { Origin of depicted specimens: USA, Alaska, Turnagin Pass, collected by Mrs. Helen Hall, August 15, 1973. } #PING. MACROCERAS SUBSP. NORTENSIS NAME: [Pinguicula macroceras subsp. nortensis] IMAGE: sj71.15.jpg DESC: sj71.15: Three views of flower (natural site specimens) IMAGE: sj71.16.jpg DESC: sj71.16: Three views of flower (natural site specimens) IMAGE: sj71.19.jpg DESC: sj71.19: Two flowers (natural site specimens), May 13, 1971 IMAGE: sj71.14.jpg DESC: sj71.14: Eight flowers (natural site specimens), May 10, 1971 IMAGE: sj71.18.jpg DESC: sj71.18: 15 calyces (natural site specimens), May 10, 1971 IMAGE: sj71.11.jpg DESC: sj71.11: Habitus (natural site specimens), May 7, 1971 IMAGE: sj71.12.jpg DESC: sj71.12: Habitus (natural site specimens), May 7, 1971 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: May 10, 1971 TEXT: { Origin of depicted specimens: USA, northernmost California, Del Norte county, Smith River South Fork (Sheep Pen Creek) between Crescent City and Gasquet, 100 m, serpentine, collected by myself May 7, 1971.
Note the different spur length in sj71.19. The short spurred specimen is likely to present what E. Hultén named P. macroceras var. microceras. I found flowers with normal and short spurs on the same plant. } #PING. MORANENSIS NAME: [Pinguicula moranensis] IMAGE: sj70.195.jpg DESC: sj70.195: Three views of flower (cultivation specimens) IMAGE: sj70.196.jpg DESC: sj70.196: Color variations (cultivation specimens), August 29, 1970 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: August 29, 1970 TEXT: { Origin of depicted specimens: University of Bern Botanical Garden. True origin unknown. } NAME: [Pinguicula moranensis and gypsicola] IMAGE: sj70.197.jpg DESC: sj70.197: Three flowers (cultivation specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: August 29, 1970 TEXT: { Large flower: P. moranensis, small flowers: P. gypsicola. } NAME: [Pinguicula moranensis] with moth IMAGE: sj94.5.21.jpg DESC: sj94.5.21: Flower with moth (cultivation specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: May 1st, 1994 TEXT: { A common moth (Plusia gamma) visiting P. moranensis. } #PING. NEVADENSIS NAME: [Pinguicula nevadensis] IMAGE: sj70.163.jpg DESC: sj70.163: Three views of flower (cultivation specimens), July 31, 1970 IMAGE: sj70.164.jpg DESC: sj70.164: Eight flowers (cultivation specimens), July 31, 1970 IMAGE: sj70.161.jpg DESC: sj70.161: Habitus (cultivation specimens), July 29, 1970 IMAGE: sj95.11.21.jpg DESC: sj95.11.21: Habitus (cultivation specimen), August 6, 1995 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 31,1970 TEXT: { Origin of depicted specimens: Spain, Prov. de Granada, Sierra Nevada, several sites below Mt. Veleta and Mt. Mulhacen summits, 2800-3000m, granite, collected by myself August 18, 1969. See also sj70.187 with P. leptoceras and other species. } #PING. RAMOSA NAME: [Pinguicula ramosa] IMAGE: sj73.19.jpg DESC: sj73.19: Flower (cultivation specimen) IMAGE: sj70.206.jpg DESC: sj70.206: Winter buds, October 12, 1970 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: April 16, 1973 TEXT: { Origin of depicted specimen: Japan, Honshu, Tochigi Pref., Mt. Nantai-san, 1900 m, volcanic substrate, collected by the late Katsuyuki Yoshimura (who was the first to determine the chromosome number of this species, October 5, 1970. Morphologically the flower of P. ramosa (2n=18) cannot be distinguished from P. variegata (2n=64) from which it differs only by the chromosomes and the stalk ramification (see sj80.49). P. ramosa is a strictly protected species! } #PING. VALLISNERIIFOLIA NAME: [Pinguicula vallisneriifolia] IMAGE: sj70.88.jpg DESC: sj70.88: Three views of flower (cultivation specimens), June 10, 1970 IMAGE: sj70.186.jpg DESC: sj70.186: Flower (cultivation specimen), June 8, 1970 IMAGE: sj94.7.17A.jpg DESC: sj94.7.17A: Flowers (cultivation specimens), May 21, 1994 IMAGE: sj94.9.14.jpg DESC: sj94.9.14: Flowers (cultivation specimens), May 28, 1994 IMAGE: sj72.20.jpg DESC: sj72.20: Habitus (cultivation specimen), June 5, 1972 IMAGE: sj72.22.jpg DESC: sj72.22: Habitus (cultivation specimen), June 5, 1972 IMAGE: sj95.4.20.jpg DESC: sj95.4.20: Spring rosettes (natural site specimens), April 16, 1995 IMAGE: sj95.4.7.jpg DESC: sj95.4.7: Habitat, April 16, 1995 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} TEXT: { Origin of depicted specimens: Spain, Prov. de Jaen, Sierra de Cazorla, 1200 m, limestone, collected by myself August 20, 1969. Grows on vertical rock walls, below overhanging rock parts in niches protected from direct rainfall. See also sj70.84 with P. alpina.
(This plant has been in cultivation since 1969!). Note the overlapping of the wide corolla lobes. The specimens from Rio Borosa (sj95.7.27) have less overlap.
Note the stolons in sj72.20 and sj72.22. After flowering Pinguicula vallisneriifolia forms the long-leaved summer rosette. Characteristically Pinguicula vallisneriifolia has also digestive glands on the underside of the leaves.
The rosettes in sj95.4.20 will soon develop flowers and stolons, followed by the long-leaved summer rosette. } NAME: [Pinguicula vallisneriifolia] IMAGE: sj95.7.27.jpg DESC: sj95.7.27: Flowers (cultivation specimens), June 4, 1995 IMAGE: sj95.7.17.jpg DESC: sj95.7.17: Habitus (cultivation specimens), June 4, 1995 IMAGE: sj95.3.23.jpg DESC: sj95.3.23: Habitat, April 16, 1995 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} TEXT: { Origin of depicted specimens: Spain, Prov. de Jaen, Sierra de Segura/Sierra del Pozo, Rio Borosa, Cerrada de Elias, 900 m, limestone, collected by myself April16, 1995. At this site the flowers are almost white and the corolla lobes are less overlapping than in the Cazorla site. } #PING. VARIEGATA NAME: [Pinguicula variegata] IMAGE: sj80.49.jpg DESC: sj80.49: Three views of flower (natural site specimens), July 21, 1980 IMAGE: sj80.50.jpg DESC: sj80.50: Three views of flower (natural site specimens), July 21, 1980 IMAGE: sj80.52.jpg DESC: sj80.52: Two flowers (natural site specimens), July 21, 1980 IMAGE: sj80.47.jpg DESC: sj80.47: Habitus (natural site), July 20, 1980 IMAGE: sj80.54.jpg DESC: sj80.54: Leaves, rosettes (natural site specimens), July 21, 1980 IMAGE: sj80.57.jpg DESC: sj80.57: Winter buds (natural site specimens), July 21, 1980 IMAGE: sj80.59.jpg DESC: sj80.59: Habitat, July 20, 1980 IMAGE: sj80.61.jpg DESC: sj80.61: Habitat, July 20, 1980 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 21, 1980 TEXT: { Origin of depicted specimens: Russia, Siberia, Baikalsky Chrjebet, Mt. Shartlay, near spring of the Lena river, 1900 m, Sphagnum bog, permafrost, collected by Vladimir Moloshnikov, Victor Voronin and myself July 20, 1980. At this site the vegetation period lasts only for 6-8 weeks, the remaining time of the year the soil is thoroughly frozen.
While flowering, the plants already develop their winter buds. They reach their full size before the seeds are ripe. On the day of this photography, July 21, the leaves of the left specimen in sj80.57 were already decaying. In mid-summer the permafrost begins 35 cm below the Sphagnum surface.
Most plants are totally overgrown by Sphagnum and peat grasses.
The scale in sj80.54 is 1 unit = 1 mm. Note the similarity with the rosettes of P. ramosa } #PING. VILLOSA NAME: [Pinguicula villosa] IMAGE: sj77.04.jpg DESC: sj77.04: Habitus (cultivation specimens) June 11, 1977 IMAGE: sj77.05.jpg DESC: sj77.05: Habitus (cultivation specimens) June 11, 1977 IMAGE: sj91.1.8A.jpg DESC: sj91.1.8A: Habitat June 15, 1991 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} TEXT: { Origin of depicted specimens: Finland, near Oulu, Sphagnum fuscum bog, collected in autumn 1976 by Pentti Martikainen. See also sj70.132 with P. grandiflora and other species.
sj91.1.8A show the habitat of P. villosa east of Oulu near Kiiminki, visited June 15, 1991. Many specimens were just in flower. } NAME: [Pinguicula villosa] IMAGE: sj81.27.jpg DESC: sj81.27: Habitus (cultivation specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 5, 1981 TEXT: { Origin of depicted specimens: USA, Alaska, Kenai island, collected August 11, 1980 by Ron Fleming. Scale unit = 1 mm. } NAME: [Pinguicula villosa] IMAGE: sj70.189.jpg DESC: sj70.189: Winter buds (natural site specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: Sept. 12, 1970 TEXT: { Origin of depicted specimens: Norway, Stabbursdalen, collected by Leif Ryvarden September 3, 1970. } NAME: [Pinguicula villosa and ramosa] IMAGE: sj70.207.jpg DESC: sj70.207: winter buds (cultivation specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: October 12, 1970 TEXT: { Top:P. villosa, bottom: P. ramosa. Scale unit = 1 mm. } #PING. VULGARIS NAME: [Pinguicula vulgaris] IMAGE: sj70.177.jpg DESC: sj70.177: Three views of flower (natural site specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: August 16, 1970 TEXT: { Origin of depicted specimens: France, Dépt. Haute-Savoye, Col des Montets/Vallée de Chamonix, 1400 m, collected by myself August 13, 1970. See also sj72.08 with P. alpina, sj70.132 with P. grandiflora, sj70.187 with P. leptoceras, sj70.146 with P. longifolia subsp. longifolia, sj70.129 with P. vulgaris f. bicolor and sj76.34 with P. vulgaris f. bicolor. } NAME: [Pinguicula vulgaris] IMAGE: sj70.130.jpg DESC: sj70.130: Three views of flower (natural site specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 11, 1970 TEXT: { Origin of depicted specimens: Switzerland, Canton Bern, Prealps, near Habkern (N of Interlaken), 1150 m, neutral soil, collected by myself July 10, 1970. } NAME: [Pinguicula vulgaris] IMAGE: sj81.18.jpg DESC: sj81.18: Three views of flower (cultivation specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: May 17, 1981 TEXT: { Origin of depicted specimens: NW Turkey, Uludag Mt. near Bursa, 2000-2200 m, collected by myself Sept. 23, 1980 . This region is indicated by S.J. Casper for P. balcanica subsp. pontica, but the plants I found are more likely to be P. vulgaris. } NAME: [Pinguicula vulgaris] IMAGE: sj72.01.jpg DESC: sj72.01: Three views of flower (cultivation specimens), May 21, 1972 IMAGE: sj72.02.jpg DESC: sj72.02: Eight flowers (cultivation specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} TEXT: { Origin of depicted specimens: USA, Michigan, upper peninsula, Pictured Rocks east of Munising (Lake Superior), 200 m, calcareous sandstone, collected by myself September 26, 1971. } NAME: [Pinguicula vulgaris (?)] IMAGE: sj74.30.jpg DESC: sj74.30: Three flowers (cultivation specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: June 2, 1974 TEXT: { Origin of depicted specimens: France, Dépt. Cantal, Le Lioran between Aurillac and Murat, , 1150 m, volcanic substrate, collected by myself August 1st, 1973. I am not sure whether this is just P. vulgaris as it has also some characteristics of P. longifolia. } NAME: [Pinguicula vulgaris] IMAGE: sj70.131.jpg DESC: sj70.131: Eight flowers (cultivation specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 11, 1970 TEXT: { Origin of depicted specimens: Switzerland, see sj70.130. Note some aberrant calyx shapes: At this site 40 percent of the plants have only 4 calyx tips (i.e. the two lower tips are fused to one). } NAME: [Pinguicula vulgaris and alpina] IMAGE: sj72.08.jpg DESC: sj72.08: Two flowers (cultivation specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: May 22, 1972 TEXT: { Origin of depicted specimens: P. vulgaris sj72.01, P. alpina sj72.06 } NAME: [Pinguicula vulgaris] IMAGE: sj72.35.jpg DESC: sj72.35: Habitus (natural site specimen) IMAGE: sj72.44.jpg DESC: sj72.44: Habitus (natural site specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 2, 1972 (72.35) and July 9, 1972 (72.44) TEXT: { Origin of depicted specimens: Switzerland, Canton Bern, Prealps, Trogenmoos/Grünenberg-Pass N of Interlaken, 1500 m, slightly acid soil. } NAME: [Pinguicula vulgaris] IMAGE: sj77.25.jpg DESC: sj77.25: Habitus (natural site specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 22, 1977 TEXT: { Origin of depicted specimens: Switzerland, Canton Ber, Prealps, Trogenalp N of Interlaken, 1700 m, slightly calcareous soil. } #PING. VULGARIS F. BICOLOR NAME: [Pinguicula vulgaris f. bicolor] IMAGE: sj70.145.jpg DESC: sj70.145: Three views of flower (cultivation specimens), July 28, 1970 IMAGE: sj70.143.jpg DESC: sj70.143: Habitus (cultivation specimen), July 26, 1970 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 28, 1970 TEXT: { Origin of depicted specimens: France, Dépt. Haute-Savoye, Solaizon above Brison (between Bonneville and Cluses), around 1500 m, limestone, collected by myself July 2, 1968. See also sj70.132 with P. grandiflora and sj70.146 with P. longifolia. } NAME: [Pinguicula vulgaris f. bicolor] IMAGE: sj70.134.jpg DESC: sj70.134: Three views of flower (cultivation specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 11, 1970 TEXT: { Origin of depicted specimens: Switzerland, Canton Vaud, Jura, nera Col du Marchairuz, 1350 m, limestone, collected by myself June 3, 1967. } NAME: [Pinguicula vulgaris with f. bicolor and f. albida] IMAGE: sj76.34.jpg DESC: sj76.34: Three flowers (cultivation specimens) June 26, 1976 IMAGE: sj70.129.jpg DESC: sj70.129: Two flowers (cultivation specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} TEXT: { In sj76.34 top: P. vulgaris, origin see sj72.35. Middle: P. vulgaris f. bicolor, origin see sj70.134. Bottom: P. vulgaris f. albida, origin sj72.35.
Origin of specimens depicted in sj70.129: Pinguicula vulgaris sj72.35, Pinguicula vulgaris f. bicolor sj70.134 } #PING. HYBRIDS NAME: [Pinguicula leptoceras x vulgaris] (or vice versa) IMAGE: sj70.167.jpg DESC: sj70.167: Three views of flower (natural site specimens), July 31, 1970 IMAGE: sj72.49.jpg DESC: sj72.49: Three views of flower (cultivation specimens), July 12, 1972 IMAGE: sj72.36.jpg DESC: sj72.36: Habitus (cultivation specimen), July 2, 1972 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 31, 1970 TEXT: { Origin of depicted specimens: Switzerland, Canton Graubuenden, eastern side of Oberalp-Pass, 2000 m, granite, collected by myself July 31, 1970. At this common site of both parent species (and in spite of the different chromosome number) hybrids are not rare. P. leptoceras begins to flower two weeks before P. vulgaris but the flowering period overlaps for about one week. } NAME: [Pinguicula grandiflora x longifolia subsp. longifolia or vice versa] IMAGE: sj81.05.jpg DESC: sj81.05: Habitus (cultivation specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: May 10, 1981 TEXT: { This is likely to be a cultivation hybrid between P. grandiflora and P. longifolia subsp. longifolia. In the Pyrenees I have seen natural hybrids between the two parent species (above Gavarnie). } NAME: [Pinguicula grandiflora x vallisneriifolia] (or vice versa) IMAGE: sj94.9.19.jpg DESC: sj94.9.19: Flowers (cultivation specimens), May 28, 1994 IMAGE: sj94.9.21.jpg DESC: sj94.9.21: Flower with pollinator (cultivation specimens), May 28, 1994 IMAGE: sj94.7.15A.jpg DESC: sj94.7.15A: Habitus (cultivation specimens), May 21, 1994 CREDIT: Juerg Steiger {steiger@iae.unibe.ch} TEXT: { A cultivation hybrid between P. grandiflora and P. vallisneriifolia caused by insects (flies, bumblebees) pollinating cultured plants. There is no common natural site of both species. } NAME: [Pinguicula longifolia(?) x vallisneriifolia] (or vice versa) IMAGE: sj94.6.10.jpg DESC: sj94.6.10: Flowers (cultivation specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: May 9, 1994 TEXT: { Cultivation hybrid caused by pollinating insects. Sometimes hybrids display characteristics which do not exist in either parent species. In the present case the hybrid shows forked spur ends which is not the case in both parent species. } NAME: [Pinguicula vallisneriifolia x vulgaris(?)] (or vice versa) IMAGE: sj95.5.26A.jpg DESC: sj95.5.26A: Habitus (cultivation specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: May 14, 1995 TEXT: { Hybrid with unclear genesis. Calyx and spur are vallisneriifolia-like, the rest of the flower is more vulgaris-like, the leaves are intermediate. } NAME: [Pinguicula grandiflora x vulgaris] IMAGE: sj94.9.30.jpg DESC: sj94.9.30: Flowers (cultivation specimens) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: May 28, 1994 TEXT: { Cultivation hybrid. I have seen this hybrid also at a natural site in the Pyrenees. It is also reported from Ireland. } NAME: [Pinguicula 'NIH 1' (not identified hybrid 1)] IMAGE: sj94.9.9.jpg DESC: sj94.9.9: Flowers (cultivation specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: May 28, 1994 TEXT: { Some characteristics of P. corsica and P. longifolia, but not clearly identifiable. } NAME: [Pinguicula 'NIH 2' (not identified hybrid 2)] IMAGE: sj94.7.21A.jpg DESC: sj94.7.21A: Habitus (cultivation specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: May 21, 1994 TEXT: { An unidentifiable hybrid. } #PINGUICULA: ABNORMALITIES NAME: [Pinguicula grandiflora] IMAGE: sj95.7a.24.jpg DESC: sj95.7a.24: Pitcher shaped and heart shaped leaf (cultivation specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: July 18, 1995 TEXT: { Pitcher shaped and heart shaped leaf (cultivation specimen) } NAME: [Pinguicula longifolia subspecies] IMAGE: sj94.10.8.jpg DESC: sj94.10.8: Pitcher shaped leaf (cultivation specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: June 4, 1994 TEXT: { Pitcher shaped leaf of Pinguicula longifolia. } NAME: [Pinguicula longifolia subsp. reichenbachiana] IMAGE: sj94.7.23A.jpg DESC: sj94.7.23A: Pitcher shaped leaf (cultivation specimen) CREDIT: Juerg Steiger {steiger@iae.unibe.ch} DATE: May 23, 1994 TEXT: { Scale unit = 1 mm }